Demonstration of how to calculate the shannon wiener diversity index in microsoft excel. Shannon weiner index the shannon index is affected by both the number of species and their equitability, or evenness. However, in practice, the difference between these two approaches is usually so small that the simpler formulas are generally acceptable magurran 2004. We will then consider one measure used to compare diversity among communities, jaccard. Pdf a simple formula for calculating the lower limit of. The geometric jensen shannon divergence or gjensen shannon divergence yields a closedform formula for gaussian distributions by taking the geometric mean. Statistics shannon wiener diversity index in the literature, the terms species richness and species diversity are sometimes used interchangeably. Species diversity concepts describing communities species. Those measures which are a cdiilbination of 5 and n total number of individuals, for instance the m2rgalefindex and the log series index a, also give these woods equal diversity table 4. The shannon diversity index h is another index that is commonly used to characterize species diversity in a community. A generalization of the jensen shannon divergence using abstract means like the geometric or harmonic means instead of the arithmetic mean was recently proposed. Putting the values into the formula for shannon index. Like simpsons index, shannon s index accounts for both abundance and evenness of the species present.
Diversity indices that combine both richness and abundance i. Dpsi log pi, where s is the total number of species in the sample, i is the total. The formulas for the true diversity of the indices are as follows. Shannon wiener species diversity index formula data analysis. If abundance is primarily concentrated into one species, the index will be close to zero. Pdf in this article, an attempt has been made to study the diversity of various religions and also. Introduction although the use of the word information, with different meanings, can be traced back to. The shannon index is an information statistic index, which means it assumes all species are.
The formula for calculating the shannon diversity index is h. Shannon wiener index of diversity information index a measure used by ecologists when a system contains too many individuals for each to be identified and examined. S, total number of species in the community richness. Equitability assumes a value between 0 and 1 with 1 being complete evenness. The shannon diversity index ranges typically from 1. Shannon index is a commonly used diversity index that takes into account.
Divide shannon s diversity index h by natural logarithm of species richness lns to calculate the species evenness. Measures of segregation and isolation benjamin forest. Index of dissimilarity d the index of dissimilarity is the most common measure of segregation. Comparing species diversity and evenness indices by c. A diversity index also called phylogenetic or simpsons diversity index is a quantitative measure that reflects how many different types such as species there are in a dataset a community and that can simultaneously take into account the phylogenetic relations among the individuals distributed among those types, such as richness, divergence or evenness.
An example of the shannon wiener diversity index for science students. How can i calculate shannons diversity index using excel. In the following we will use the shannon diversity index h in other applications also named shannon entropy which is based on the geometric mean, and the true diversity of order one. Note that species evenness ranges from zero to one, with zero signifying. I am in need of calculating shannon s diversity index of my identified diatom species using excel with the formula h. Simpsons index d is a measure of diversity, which takes into account both species richness, and an evenness of abundance among the species present. Pdf a simple formula for calculating the lower limit of shannons.
The actual formula for the shannon weiner diversity index is. If f2l 1r and f, the fourier transform of f, is supported. It turns out that the mathematical relationships hold true whether one is dealing with molecules in solution or species in an ecological community. The formulas given in table table1 1 are for complete populations, and the actual formulas for calculating these indices from sample data are slightly more complex magurran 2004. The maximum diversity hmax of a sample is found when all species are equally abundant. Learn how to calculate shannon wiener diversity index with clear definition, formula and example. This comparison can be between two different habitats or a comparison of one habitat over time. It uses the logarithm, and we will write it here with the natural logarithm. Historically, the index has been used to measure the effects of habitat quality such as effects of pollution effluents. A measure used by ecologists when a system contains too many individuals for each to be identified and examined. Engels department of zoology, state university of ghent, belgium in a low diversity brackish water habitat the diversity of the copepod community is best measured with the shannon wiener information function and its evenness by an index proposed by the first author eh181.
A greater number of species and a more even distribution both increase diversity as measured by h. The index of dissimilarity for two groups, whites and blacks, in a particular city. The shannon wiener diversity index is one measure that we will use to try to draw information from samples in the field. To understand dembski, you need a little background on shannon. Where, h shannon index of diversity pi the proportion of important value of the i. Pdf application of shannons index to study diversity with. The variance of is calculated by and a statistic to test the significant differences between two plots or samples as where is the shannon diversity index of sample. How can i calculate shannon s diversity index using excel sheet. The simpson index is a dominance index because it gives more weight to common or dominant species. In essence it measures the probability that two individuals randomly selected from an area will belong to the same species. Shannon s diversity index and simpsons index of dominance were calculated using important value index ivi of species.
Although it has limitations, it is relatively easy to calculate and to interpret. According to local occurrence status, there were 12 species ranked as very abundant, 19 abundant, 62 very. The shannon index is a measurement used to compare diversity between habitat samples. A diversity index is the measure of species diversity in a given community. A simple formula for calculating the lower limit of shannon s diversity index article pdf available in ecological modelling 992. When all species in the data set are equally common, all pi values 1r and the shannon weiner index equals lnr. The shannon sampling theorem and its implications gilad lerman notes for math 5467 1 formulation and first proof the sampling theorem of bandlimited functions, which is often named after shannon, actually predates shannon 2. Application of shannons index to study diversity with.
The shannon diversity index h is another index that is commonly used to. Simpsons index of diversity in ecology, this index is used to quantify the biodiversity of a habitat. A a diversity index is a mathematical measure of species diversity in a given community. Statistics shannon wiener diversity index tutorialspoint. The methods on offer range from the familiar, such as the calculation of. How to calculate shannon wiener diversity index formula. Shannon and simpson values for 1987 and 1992 using the alpha diversity tab. Pdf the purpose of this note is to demonstrate that the exact value of the lower limit of shannons diversity index called hmin. Shannons diversity index real statistics using excel. It is different from species richness in that unlike richness it also shows community composition and takes into account the relative abundance of species that are present in the community. A simple formula for calculating the lower limit of shannons diversity.
The more unequal the abundance of species, the larger the weighted geometric mean of the pi values, the smaller the index. In particular, for a random sample, we can use shannon s index of diversity aka as shannon weiners index, which is defined as. For the following sections, s represents the number of species. H sumpi lnpi ehh max where, sum summation pi numbe of individuals of species itotal number of samples s number of species or species richness h max maximum diversity possible e evenesshh max. In the shannon index, p is the proportion nn of individuals of one particular species found n divided by the total number of individuals found n, ln is the natural log.
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